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Event ID 4999 and Event ID 7031 are logged when you move a mailbox to an Exchange Server 2010 server
Article ID: 979862 - View products that this article applies to.
When you try to move a mailbox from an earlier version of Exchange server to a Microsoft Exchange Server 2010 server, the operation fails. Additionally, you find the following events are logged on the Exchange Server 2010 server.
This problem occurs because a message in the effected mailbox misses a certain property.
To resolve this problem, install the following update rollup:
(http://support.microsoft.com/kb/982639/ )Description of Update Rollup 4 for Exchange Server 2010
Microsoft has confirmed that this is a problem in the Microsoft products that are listed in the "Applies to" section.